Scottish soldier Donnie MacRae died as a German prisoner of war during World War Two - but it was not until almost 80 years later that his family discovered he had been buried without his brain.

Donnie died in a PoW hospital in 1941 and because he had suffered with a rare neurological condition an autopsy was performed on his body.

During the post-mortem, his brain and part of his spinal cord were removed and sent to the Kaiser Wilhelm Institute for Psychiatry in Munich to be used for research.

His body was buried by the Germans and later reburied by the Allies in the Commonwealth War Graves cemetery in Berlin but no-one knew his brain had been removed.

In total, about 160 small slices of Donnie’s brain and spinal cord have been kept in the archives of the Munich research centre - since renamed the Max Planck Institute for Psychiatry - ever since.

A BBC Radio 4 documentary - Shadow of War: A Tainted Anatomy - looks at why this happened and at the work being carried out to reunite the remains with the soldier in his grave.